.species-page,
.species-directory-page {
  max-width: 1180px;
  margin: 0 auto;
}

.species-breadcrumb {
  display: flex;
  gap: 10px;
  align-items: center;
  color: var(--muted-strong, #42524b);
  font-size: 13px;
  margin-bottom: 48px;
}

.species-breadcrumb a {
  color: inherit;
  text-decoration: none;
}

.species-breadcrumb a:hover,
.species-breadcrumb a:focus-visible {
  color: var(--accent, #0c7b63);
}

.species-hero,
.species-directory-hero {
  max-width: 820px;
  padding: 22px 0 76px;
}

.species-hero h1,
.species-directory-hero h1 {
  margin: 0;
  font-size: clamp(48px, 8vw, 92px);
  line-height: .98;
  letter-spacing: -.04em;
}

.species-scientific {
  margin: 18px 0 0;
  color: var(--muted-strong, #42524b);
  font-size: 17px;
}

.species-hero .lead,
.species-directory-hero .lead {
  max-width: 650px;
  margin-top: 28px;
}

.species-art {
  max-width: 360px;
  margin: 28px 0 8px;
}

.species-art img {
  display: block;
  width: 100%;
  aspect-ratio: 1.94;
  object-fit: cover;
  border-radius: 12px;
  background: #e8f0e8;
}

.species-art figcaption {
  margin-top: 10px;
  color: var(--muted-strong, #42524b);
  font-size: 14px;
  line-height: 1.6;
}

.species-alias {
  color: var(--muted-strong, #42524b);
  font-size: 14px;
}

.species-status {
  display: inline-flex;
  gap: 14px;
  align-items: baseline;
  margin-top: 26px;
  padding: 11px 14px;
  border-left: 2px solid var(--accent, #0c7b63);
  background: var(--accent-soft, rgba(12, 123, 99, .12));
}

.species-status span {
  font-size: 14px;
}

.species-status small {
  color: var(--muted-strong, #42524b);
  font-size: 12px;
}

.species-grid,
.species-stage-grid {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 16px;
}

.species-intro article,
.species-stage,
.species-facts,
.species-record,
.species-cta,
.species-directory-note {
  border-top: 1px solid var(--line, rgba(23, 38, 32, .15));
}

.species-intro article {
  padding: 28px 0 34px;
}

.species-intro h2,
.species-stage h3 {
  margin: 0 0 12px;
  font-size: 20px;
  line-height: 1.22;
}

.species-intro p:last-child,
.species-stage p,
.species-facts dd,
.species-record p,
.species-cta p {
  color: var(--muted-strong, #42524b);
  line-height: 1.75;
}

.species-facts {
  margin-top: 76px;
  padding: 58px 0;
}

.species-facts dl {
  display: grid;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  gap: 0 40px;
  margin: 34px 0 0;
}

.species-facts dl > div {
  padding: 22px 0;
  border-top: 1px solid var(--line, rgba(23, 38, 32, .15));
}

.species-facts dt {
  font-size: 13px;
  color: var(--accent, #0c7b63);
}

.species-facts dd {
  margin: 8px 0 0;
}

.species-stages {
  padding: 58px 0;
}

.species-stage {
  padding: 28px 0 18px;
}

.species-stage ul {
  padding-left: 18px;
  color: var(--muted-strong, #42524b);
  line-height: 1.8;
}

.species-record {
  display: grid;
  grid-template-columns: 1.2fr .8fr;
  gap: 60px;
  padding: 58px 0;
}

.species-record h2,
.species-cta h2 {
  margin: 0;
  font-size: clamp(28px, 4vw, 46px);
  line-height: 1.1;
}

.species-boundary {
  align-self: end;
  padding-left: 24px;
  border-left: 1px solid var(--accent, #0c7b63);
}

.species-cta {
  padding: 58px 0 80px;
}

.species-cta .button {
  margin: 16px 18px 0 0;
}

.species-cta .text-link {
  display: inline-block;
  margin-top: 20px;
}

.species-directory-group {
  padding: 44px 0 12px;
}

.species-directory-grid {
  display: grid;
  grid-template-columns: repeat(4, minmax(0, 1fr));
  gap: 0;
  margin-top: 18px;
  border-top: 1px solid var(--line, rgba(23, 38, 32, .15));
  border-left: 1px solid var(--line, rgba(23, 38, 32, .15));
  background: transparent;
}

.species-directory-grid a {
  display: flex;
  min-height: 86px;
  flex-direction: column;
  justify-content: space-between;
  padding: 18px;
  border-right: 1px solid var(--line, rgba(23, 38, 32, .15));
  border-bottom: 1px solid var(--line, rgba(23, 38, 32, .15));
  color: inherit;
  text-decoration: none;
  background: var(--panel, #fbfcf8);
  transition: background .18s ease, color .18s ease;
}

.species-directory-grid a:hover,
.species-directory-grid a:focus-visible {
  color: var(--accent-ink, #fff);
  background: var(--accent, #0c7b63);
}

.species-directory-grid small {
  color: var(--muted-strong, #42524b);
  font-size: 12px;
}

.species-directory-grid em {
  display: -webkit-box;
  min-height: calc(1.55em * 3);
  max-height: calc(1.55em * 3);
  overflow: hidden;
  margin-top: 8px;
  color: var(--muted-strong, #42524b);
  font-size: 11.5px;
  font-style: normal;
  line-height: 1.55;
  -webkit-box-orient: vertical;
  -webkit-line-clamp: 3;
}

.species-directory-grid a:hover em,
.species-directory-grid a:focus-visible em {
  color: inherit;
}

.species-directory-grid a:hover small,
.species-directory-grid a:focus-visible small {
  color: inherit;
}

.species-directory-note {
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 28px;
  margin-top: 70px;
  padding: 28px 0 80px;
}

.species-directory-note p {
  max-width: 720px;
  margin: 0;
  color: var(--muted-strong, #42524b);
  line-height: 1.7;
}

@media (max-width: 780px) {
  .species-grid,
  .species-stage-grid,
  .species-facts dl,
  .species-record {
    grid-template-columns: 1fr;
  }

  .species-directory-grid {
    grid-template-columns: repeat(2, minmax(0, 1fr));
  }

  .species-status,
  .species-directory-note {
    align-items: flex-start;
    flex-direction: column;
  }
}